Journal Articles

Publications are in reverse chronological order. Within the same year, they are sorted by the last name of the first author, then by number of authors, then by last name of the second author. Send me an email if you would like to request a PDF.

Since 2020


  • Eldridge, M.D.B, R.M.D. Beck, D.A. Croft, K.J. Travouillon & B.J. Fox. 2019. An emerging consensus in the evolution, phylogeny and systematics of marsupials and their fossil relatives (Metatheria). Journal of Mammalogy 100(3):802–837.
  • Engelman, R.K. & D.A. Croft. 2019. Strangers in a strange land: Ecological dissimilarity to metatherian carnivores may partly explain early colonization of South America by Cyonasua-group procyonids (Carnivora). Paleobiology 45:598-611.
  • Brandoni, D., A.A. Carlini, F. Anaya, P. Gans & D.A. Croft. 2018. New remains of Megathericulus patagonicus Ameghino (Xenarthra, Tardigrada) from the Serravallian (middle Miocene) of Bolivia; chronologic and biogeographic implications. Journal of Mammalian Evolution 25:327-337. 
  • Croft, D.A., R.K. Engelman, T. Dolgushina & G. Wesley. 2018. Diversity and disparity of sparassodonts (Metatheria) reveal non-analogue nature of ancient South American mammalian carnivore guilds. Proceedings of the Royal Society B: Biological Sciences 285:20172012.
  • Engelman, R.K., J.J. Flynn, P. Gans, A.R. Wyss, & D.A. Croft. 2018. Chlorocyon phantasma, a late Eocene borhyaenoid (Mammalia: Metatheria: Sparassodonta) from the Los Helados locality, Andean Main Range, central Chile. American Museum Novitates 3918:1-22.
  • McGrath, A.J., F. Anaya & D.A. Croft. 2018. Two new macraucheniids (Mammalia: Litopterna) from the late middle Miocene (Laventan South American Land Mammal Age) of Quebrada Honda, Bolivia. Journal of Vertebrate Paleontology 38(3):e1461632.
  • Wyss, A.R., J.J. Flynn & D.A. Croft. 2018. New Paleogene notohippids and leontiniids (Toxodontia; Notoungulata; Mammalia) from the early Oligocene Tinguiririca Fauna of the Andean Main Range, Central Chile. American Museum Novitates 3903:1-42.
  • Arnal, M., M.G. Vucetich, D.A. Croft, M.S. Bargo, J.C. Fernicola & S.F. Vizcaíno. 2017. Systematic revision and evolutionary history of Acarechimys Patterson in Kraglievich 1965 (Rodentia, Caviomorpha, Octodontoidea). Ameghiniana 54(3):307–330.
  • Catena, A.M., D.I. Hembree, B.Z. Saylor, F. Anaya & D.A. Croft. 2017. Paleosol and ichnofossil evidence for significant Neotropical habitat variation during the late middle Miocene (Serravallian). Palaeogeography, Palaeoclimatology, Palaeoecology 487:381-398.
  • Engelman, R.K., F. Anaya & D.A. Croft. 2017. New palaeothentid marsupials (Paucituberculata) from the middle Miocene of Quebrada Honda, Bolivia, and their implications for the palaeoecology, decline and extinction of the Palaeothentoidea. Journal of Systematic Palaeontology 15: 787-820.
  • Antoine, P-O., M.A. Abello, S. Adnet, A.J. Altamira, P. Baby, G. Billet, M. Boivin, Y. Calderón, A. Candela, J. Chabin, F. Corfu, D.A. Croft, M. Ganerød, C. Jaramillo, S. Klaus, L. Marivaux, R. Navarrete, M.J. Orliac, F. Parra, M.E. Pérez, F. Pujos, J.-C. Rage, A. Ravel, C. Robinet, M. Roddaz, J.V. Tejada-Lara, J. Vélez-Juarbe, F. Wesselingh & R. Salas Gismondi. 2016. A 60-million-year Cenozoic history of western Amazonian ecosystems in Contamana, eastern Peru. Gondwana Research 31:30-59.
  • Catena, A.M., D.I. Hembree, B.Z. Saylor, F. Anaya & D.A. Croft. 2016. Paleoenvironmental analysis of the Neotropical fossil mammal site of Cerdas, Bolivia (middle Miocene) based on ichnofossils and paleopedology. Palaeogeography, Palaeoclimatology, Palaeoecology 459:423-439
  • Croft, D.A., A.A. Carlini, M.R. Ciancio, D. Brandoni, N.E. Drew, R.K. Engelman & F. Anaya. 2016. New mammal faunal data from Cerdas, Bolivia, a middle-latitude Neotropical site that chronicles the end of the Middle Miocene Climatic Optimum in South America. Journal of Vertebrate Paleontology 36(5):e1163574.
  • Engelman, R.K. & D.A. Croft. 2016. Evidence for sexual dimorphism in the early Miocene palaeothentid Acdestis owenii (Marsupialia: Paucituberculata). Ameghiniana 53(4):444-462.
  • Bradham, J., J.J. Flynn, D.A. Croft & A.R. Wyss. 2015. New notoungulates (Notostylopidae and basal toxodontians) from the early Oligocene Tinguiririca Fauna of the Andean Main Range, central Chile. American Museum Novitates 3841:1-24.
  • Cadena, E.A., Anaya, F. & D.A. Croft. 2015. Giant fossil tortoise and freshwater chelid turtle remains from the middle Miocene, Quebrada Honda, Bolivia: evidence for lower paleoelevations for the southern Altiplano. Journal of South American Earth Sciences 64:190-198.
  • Engelman, R.K., F. Anaya & D.A. Croft. 2015. New specimens of Acyon myctoderos (Metatheria, Sparassodonta) from Quebrada Honda, Bolivia. Ameghiniana 52(2):204-225.
  • Gaudin, T.J. & D.A. Croft. 2015. Paleogene Xenarthra and the evolution of South American mammals. Journal of Mammalogy 96(4):622-634.


  • Engelman, R.K. & D.A. Croft. 2014. A new species of small-bodied sparassodont (Mammalia: Metatheria) from the middle Miocene locality of Quebrada Honda, Bolivia. Journal of Vertebrate Paleontology 34(3):672-688.
  • Croft, D.A. 2013. What constitutes a fossil mammal community in the early Miocene Santa Cruz Formation? Journal of Vertebrate Paleontology 33(2):401-409.
  • Macrini, T.E., J.J. Flynn, X. Ni, D.A. Croft & A.R. Wyss. 2013. Comparative study of notoungulate (Placentalia, Mammalia) bony labyrinths and new phylogenetically informative inner ear characters. Journal of Anatomy 223(5):442-461.
  • Antoine, P-O., L. Marivaux, D.A. Croft, G. Billet, M. Ganerød, C. Jaramillo, T. Martin, M.J. Orliac, J. Tejada, A.J. Altamirano, F. Duranthon, G. Fanjat, S. Rousse & R. Salas Gismondi. 2012. Middle Eocene rodents from Peruvian Amazonia reveal the pattern and timing of caviomorph origins and biogeography. Proceedings of the Royal Society B: Biological Sciences 279:1319-1326.
  • Bertrand, O.C., J.J. Flynn, D.A. Croft & A.R. Wyss. 2012. Two new taxa (Caviomorpha, Rodentia) from the early Oligocene Tinguiririca Fauna, (Chile). American Museum Novitates 3750:1-36.
  • Shockey, B.J., J.J. Flynn, D.A. Croft, P. Gans & A.R. Wyss. 2012. New leontiniid Notoungulata (Mammalia) from Chile and Argentina: comparative anatomy, character analysis, and phylogenetic hypotheses. American Museum Novitates 3737:1-64.
  • Croft, D.A., J.M.H. Chick & F. Anaya. 2011. New middle Miocene caviomorph rodents from Quebrada Honda, Bolivia. Journal of Mammalian Evolution 18(4):245-268.
  • Croft, D.A., K. Niemi & A. Franco. 2011. Incisor morphology reflects diet in caviomorph rodents. Journal of Mammalogy 92(4):871-879.
  • Macrini, T.E., J.J. Flynn, D.A. Croft & A.R. Wyss. 2010. Inner ear of a notoungulate placental mammal: anatomical description and examination of potentially phylogenetically informative characters. Journal of Anatomy 216(5):600-610.
  • Townsend, B. & D.A. Croft. 2010. Middle Miocene mesotheriine diversity at Cerdas, Bolivia, and a reconsideration of Plesiotypotherium minus. Palaeontologia Electronica 13.1.1A:1-36.


  • Croft, D.A., F. Anaya, D. Auerbach, C. Garzione & B.J. MacFadden. 2009. New data on Miocene Neotropical provinciality from Cerdas, Bolivia. Journal of Mammalian Evolution 16(3):175-198.
  • Croft, D.A. & L.C. Anderson. 2008. Locomotion in the extinct notoungulate Protypotherium. Palaeontologia Electronica 11.1.1A: 1-20.
  • Croft, D.A. & D. Weinstein. 2008. The first application of the mesowear method to endemic South American ungulates (Notoungulata). Palaeogeography, Palaeoclimatology, Palaeoecology 269(1-2):103-114.
  • Croft, D.A., J.J. Flynn & A.R. Wyss. 2008. The Tinguiririca Fauna of Chile and the early stages of “modernization” of South American mammal faunas. Arquivos do Museu Nacional 66(1):191-211.
  • Flynn, J.J., R. Charrier, D.A. Croft, P.B. Gans, T.M. Heriott, J.A. Wertheim & A.R. Wyss. 2008. Chronologic implications of new Miocene mammals from the Cura-Mallín and Trapa Trapa formations, Laguna del Laja region, south central Chile. Journal of South American Earth Sciences 26(4):412-423.
  • Reguero, M.A., D.A. Croft, G.M. López & R.N. Alonso. 2008. Eocene archaeohyracids (Mammalia: Notoungulata: Hegetotheria) from the Puna, northwest Argentina. Journal of South American Earth Sciences 26(2)225-233.
  • Townsend, K.E. & D.A. Croft. 2008. Diets of notoungulates from the Santa Cruz Formation, Argentina: new evidence from enamel microwear. Journal of Vertebrate Paleontology 28(1):217-230.
  • Townsend, K.E. & D.A. Croft. 2008. Enamel microwear in caviomorph rodents. Journal of Mammalogy 89(3):729-742.
  • Croft, D.A. 2007. The middle Miocene (Laventan) Quebrada Honda Fauna, southern Bolivia, and a description of its notoungulates. Palaeontology 50(1):277-303.
  • Croft, D.A., J.J. Flynn, & A.R. Wyss. 2007. A new basal glyptodontid and other Xenarthra of the early Miocene Chucal Fauna, northern Chile. Journal of Vertebrate Paleontology 27(4):781-797. [Featured article]
  • Shockey, B.J., D.A. Croft & F. Anaya. 2007. Analysis of function in the absence of extant functional homologues: a case study using mesotheriid notoungulates (Mammalia). Paleobiology 33(2):227-247.
  • Croft, D.A. 2006. Do marsupials make good predators? Insights from predator-prey diversity ratios. Evolutionary Ecology Research 8(7):1193-1214. [Featured on cover]
  • Croft, D.A. & F. Anaya. 2006. A new middle Miocene hegetotheriid (Notoungulata: Typotheria) and a phylogeny of the Hegetotheriidae. Journal of Vertebrate Paleontology 26(2):387-399.
  • Croft, D.A., L.R. Heaney, J.J. Flynn & A.P. Bautista. 2006. Fossil remains of a new, diminutive Bubalus (Artiodactyla: Bovidae: Bovini) from Cebu Island, Philippines. Journal of Mammalogy 87(5):1037-1051. [Featured on BioOne]
  • Silverstein, J.C., J.M. Ehrenfeld, D.A. Croft, F.W. Dech, S. Small & S. Cook. 2006. Tele-immersion: preferred infrastructure for anatomy instruction. Journal of Computing in Higher Education 18(1):80-93.
  • Charrier, R., A. Chávez, S. Elgueta, G. Hérail, J.J. Flynn, D.A. Croft, A.R. Wyss & M. García. 2005. Rapid tectonic and paleogeographic evolution associated with the development of the Chucal anticline and the Chucal-Lauca Basin in the Altiplano of Arica, northern Chile. Journal of South American Earth Sciences 19(1):35-54.
  • Flynn, J.J., D.A. Croft, R. Charrier, A.R. Wyss, G. Hérail & M. García. 2005. New Mesotheriidae (Mammalia, Notoungulata, Typotheria), geochronology and tectonics of the Caragua area, northernmost Chile. Journal of South American Earth Sciences 19(1):55-74.
  • Woodman, N. & D.A. Croft. 2005. Fossil shrews from Honduras: significance for evolution in Cryptotis (Mammalia: Soricomorpha: Soricidae). Fieldiana: Geology(New Series) 51:1-30.

prior to 2005

  • Croft, D.A., J.J. Flynn & A.R. Wyss. 2004. Notoungulata and Litopterna of the early Miocene Chucal Fauna, northern Chile. Fieldiana: Geology (New Series) 50:1-49.
  • Croft, D.A., T. Kaye, & L. Panko. 2004. A new method for finding small vertebrate fossils: ultraviolet light at night. Palaeontology 47(4):795-800.
  • Croft, D.A., M. Bond, J.J. Flynn, M. Reguero & A.R. Wyss. 2003. Large archaeohyracids (Typotheria, Notoungulata) from central Chile and Patagonia including a revision of Archaeotypotherium. Fieldiana: Geology (New Series) 49:1-38.
  • Croft, D.A., J.P. Radic, E. Zurita, R. Charrier, J.J. Flynn & A.R. Wyss. 2003. A Miocene toxodontid (Mammalia: Notoungulata) from the sedimentary series of the Cura-Mallín Formation, Lonquimay, Chile. Revista Geológica de Chile 30(2):285-298.
  • Flynn, J.J, A.R. Wyss, D.A. Croft, & R. Charrier. 2003. The Tinguiririca Fauna, Chile: biochronology, paleoecology, biogeography, and a new earliest Oligocene South American land mammal “age”. Palaeogeography, Palaeoclimatology, Palaeoecology 195(3-4):229-259.
  • Reguero, M., D.A. Croft, J.J. Flynn, & A.R. Wyss. 2003. Small archaeohyracids (Typotheria, Notoungulata) from Chubut, Argentina and central Chile: trans-Andean temporal correlation. Fieldiana: Geology (New Series) 48:1-17.
  • Flynn, J.J., D.A. Croft, R. Charrier, G. Hérail & A.R. Wyss. 2002. The first Cenozoic mammal fauna from the Chilean Altiplano. Journal of Vertebrate Paleontology 22(1):200-206.
  • Croft, D.A. 2001. Cenozoic environmental change in South America as indicated by mammalian body size distributions (cenograms). Diversity and Distributions 7(6):271-287.
  • Croft, D.A. & H.A. Semken, Jr. 1994. Distribution of mammalian osteological elements recovered from waterscreened features, house fill, and overburden of the Wall Ridge Earthlodge (13ML176), Mills County, Iowa. Current Research in the Pleistocene 11:65-67.